We show that averaged sex-biased expression changes accumulate monotonically over time within the genus. simulans strains; b, the melanogaster subgroup (D.melanogaster, D. In b and c, the species A and B designation is arbitrary, but A is assigned to the species in the pair most closely related to D. d, Neighbour-joining trees with branch lengths inferred using sequence distance (genomic mutation distance There was an especially clear relationship between sequence and expression divergence.
melanogaster adults is substantial, primarily owing to gametogenesis. Generally, we observed greater male-biased expression (~7–14% of the transcriptome) relative to female-biased expression (~3–9% of the transcriptome), at a significance value of P0.01 (Mann–Whitney, false-discovery-rate-corrected). pseudoobscura (~16% female- and male-biased expression) and D. b, Expression ratio standard deviations among members of the melanogaster subgroup (D. We found no significant correlation between orthologue expression divergence between groups of species (r = 0.08, Fig. Genes with greater expression divergence in the melanogaster subgroup and the remaining species are different.
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It is, however, a native or 'first' lan- guage only in a nnitp restricted area consisting of the islands of Zanzibar and Pemba and the oppo- site coast, roughly from Dar es Salaam to Mombasa, Outside this relatively small territory, elsewhere in Kenya, in Tanzania (formerly Tanganyika), and to a lesser degree in Uganda, in the Republic of the Congo, and in other fringe regions hard to delimit, Swahili is a lingua franca of long standing, a 'second' (or 'third' or 'fourth') language enjoy- ing a reasonably well accepted status as a supra-tribal or supra-regional medium of communication.
Its chief competitor in this function is English, and English is of course a formidable competi- tor indeed in Kenya and Tanzania and particularly in Uganda.
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